Holles Natalie C. 4

4 · Aura Biosciences, Inc. · Filed May 19, 2026

AI-generated summary of this filing

Updated May 19, 2026

Aura Biosciences CEO Natalie Holles Receives Stock Awards

What Happened

Natalie C. Holles, Chief Executive Officer and President of Aura Biosciences (AURA), was granted a total of 3,323,065 award units on May 19, 2026: 600,118 restricted stock units (RSUs), 553,844 performance RSUs (PRSUs), and 2,169,103 derivative shares (option-type award). All grants were reported as awards (code A) at $0.00 per share (no cash paid; grant value reported as $0).

Key Details

Transaction date: 2026-05-19; filing date: 2026-05-19 (filed same day).
Award breakdown: 600,118 RSUs (F1); 553,844 PRSUs (F2); 2,169,103 derivative/option-style award (F3).
Vesting highlights:
- RSUs (F1): Vest in four substantially equal annual installments beginning April 15, 2027, subject to continued service.
- PRSUs (F2): Subject to time-based vesting (four equal annual installments starting April 15, 2027) and a performance condition; expire on the earlier of the 6th anniversary or termination of service if unvested.
- Derivative/option (F3): 25% of underlying shares vest April 30, 2027; remaining shares vest pro rata in 36 monthly installments thereafter, subject to continued service.
Shares owned after transaction: Not specified in the filing.
These awards were granted as inducement awards (to induce service), per footnotes.

Context

These are grants (not open-market purchases or sales). RSUs and PRSUs convert to stock only if vesting conditions are met; PRSUs additionally require achievement of performance goals. The derivative award appears to be option-like with time-based vesting; no immediate sale or cash exercise occurred. Inducement grants are common for new hires or role changes and do not, by themselves, indicate a buy/sell market signal.

Form 4

Period: 2026-05-19

DirectorSee Remarks

Transactions

Award
Common Stock
[F1]
2026-05-19+600,118→ 600,118 total
Award
Common Stock
[F2]
2026-05-19+553,844→ 1,153,962 total
Award
Stock Option (Right to Buy)
[F3]
2026-05-19+2,169,103→ 2,169,103 total
Exercise: $7.39Exp: 2036-05-19→ Common Stock (2,169,103 underlying)

Footnotes (3)

[F1]These shares were acquired pursuant to a restricted stock unit ("RSU") award as Inducement Awards. Each RSU represents the right to receive one share of the Issuer's common stock. These RSUs vest in four substantially equal annual installments beginning on April 15, 2027, subject to the Reporting Person's continued service as of each such vesting date.
[F2]These shares were acquired pursuant to a performance RSU ("PRSU") award as Inducement Awards. The PRSUs are subject to both time-based vesting and the achievement of a performance condition, both of which must be satisfied before the PRSUs will be deemed vested. The PRSUs shall vest in four substantially equal annual installments commencing on April 15, 2027, subject to Ms. Holles' continued service as of each such time-based vesting date and the satisfaction of the performance condition. The expiration date of the PRSUs is the earlier of (i) the sixth (6th) anniversary of the grant date and (ii) the date Ms. Holles no longer has a service relationship with the Issuer. Any such PRSUs that have not vested on or prior to such expiration date shall be forfeited for no consideration.
[F3]The shares underlying this option vest as follows: 25% of the shares vest on April 30, 2027 with the remainder vesting thereafter pro-rata in 36 monthly installments, subject to the Reporting Person's continued service as of each such vesting date.

Signature

/s/ Conor Kilroy, as Attorney-in-Fact|2026-05-19

1 file